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Cell culture and treatment Human bronchial epithelial cells (HBECs) were cultured in DMEM (Hyclone, Logan, UT, USA) added with 10% fetal bovine serum (Gibco, USA), 100 U/mL penicillin and 100 U/mL streptomycin (Thermo Fisher Scientific, Waltham, MA, USA) at 37C in a 5% CO 2 humidified incubator

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